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Topic Review
Permian–Triassic Extinction Event
The Permian–Triassic extinction event, also known as the P–Tr extinction, the P–T extinction, the End-Permian Extinction, and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as well as between the Paleozoic and Mesozoic eras, approximately 252 million years ago. It is the Earth's most severe known extinction event, with up to 96% of all marine species and 70% of terrestrial vertebrate species becoming extinct. It was the largest known mass extinction of insects. Some 57% of all biological families and 83% of all genera became extinct. There is evidence for one to three distinct pulses, or phases, of extinction. Potential causes for those pulses include one or more large meteor impact events, massive volcanic eruptions (such as the Siberian Traps), and climate change brought on by large releases of underwater methane or methane-producing microbes. The speed of the recovery from the extinction is disputed. Some scientists estimate that it took 10 million years (until the Middle Triassic), due both to the severity of the extinction and because grim conditions returned periodically for another 5 million years. However, studies in Bear Lake County, near Paris, Idaho, showed a relatively quick rebound in a localized Early Triassic marine ecosystem, taking around 2 million years to recover, suggesting that the impact of the extinction may have been felt less severely in some areas than others.
  • 3.4K
  • 11 Oct 2022
Topic Review
History of Life
The history of life on Earth traces the processes by which living and fossil organisms evolved, from the earliest emergence of life to the present day. Earth formed about 4.5 billion years ago (abbreviated as Ga, for gigaannum) and evidence suggests that life emerged prior to 3.7 Ga. Although there is some evidence of life as early as 4.1 to 4.28 Ga, it remains controversial due to the possible non-biological formation of the purported fossils. The similarities among all known present-day species indicate that they have diverged through the process of evolution from a common ancestor. Only a very small percentage of species have been identified: one estimate claims that Earth may have 1 trillion species. However, only 1.75–1.8 million have been named and 1.8 million documented in a central database. These currently living species represent less than one percent of all species that have ever lived on Earth. The earliest evidence of life comes from biogenic carbon signatures and stromatolite fossils discovered in 3.7 billion-year-old metasedimentary rocks from western Greenland. In 2015, possible "remains of biotic life" were found in 4.1 billion-year-old rocks in Western Australia. In March 2017, putative evidence of possibly the oldest forms of life on Earth was reported in the form of fossilized microorganisms discovered in hydrothermal vent precipitates in the Nuvvuagittuq Belt of Quebec, Canada, that may have lived as early as 4.28 billion years ago, not long after the oceans formed 4.4 billion years ago, and not long after the formation of the Earth 4.54 billion years ago. Microbial mats of coexisting bacteria and archaea were the dominant form of life in the early Archean Epoch and many of the major steps in early evolution are thought to have taken place in this environment. The evolution of photosynthesis, around 3.5 Ga, eventually led to a buildup of its waste product, oxygen, in the atmosphere, leading to the great oxygenation event, beginning around 2.4 Ga. The earliest evidence of eukaryotes (complex cells with organelles) dates from 1.85 Ga, and while they may have been present earlier, their diversification accelerated when they started using oxygen in their metabolism. Later, around 1.7 Ga, multicellular organisms began to appear, with differentiated cells performing specialised functions. Sexual reproduction, which involves the fusion of male and female reproductive cells (gametes) to create a zygote in a process called fertilization is, in contrast to asexual reproduction, the primary method of reproduction for the vast majority of macroscopic organisms, including almost all eukaryotes (which includes animals and plants). However the origin and evolution of sexual reproduction remain a puzzle for biologists though it did evolve from a common ancestor that was a single celled eukaryotic species. Bilateria, animals having a left and a right side that are mirror images of each other, appeared by 555 Ma (million years ago). Algae-like multicellular land plants are dated back even to about 1 billion years ago, although evidence suggests that microorganisms formed the earliest terrestrial ecosystems, at least 2.7 Ga. Microorganisms are thought to have paved the way for the inception of land plants in the Ordovician period. Land plants were so successful that they are thought to have contributed to the Late Devonian extinction event. (The long causal chain implied seems to involve the success of early tree archaeopteris (1) drew down CO2 levels, leading to global cooling and lowered sea levels, (2) roots of archeopteris fostered soil development which increased rock weathering, and the subsequent nutrient run-off may have triggered algal blooms resulting in anoxic events which caused marine-life die-offs. Marine species were the primary victims of the Late Devonian extinction.) Ediacara biota appear during the Ediacaran period, while vertebrates, along with most other modern phyla originated about 525 Ma during the Cambrian explosion. During the Permian period, synapsids, including the ancestors of mammals, dominated the land, but most of this group became extinct in the Permian–Triassic extinction event 252 Ma. During the recovery from this catastrophe, archosaurs became the most abundant land vertebrates; one archosaur group, the dinosaurs, dominated the Jurassic and Cretaceous periods. After the Cretaceous–Paleogene extinction event 66 Ma killed off the non-avian dinosaurs, mammals increased rapidly in size and diversity. Such mass extinctions may have accelerated evolution by providing opportunities for new groups of organisms to diversify.
  • 2.8K
  • 18 Oct 2022
Topic Review
Gorgonopsia
Gorgonopsia (from the Greek Gorgon, a mythological beast, and óps 'aspect') is an extinct clade of sabre-toothed therapsids from the Middle to Upper Permian roughly 265 to 252 million years ago. They are characterised by a long and narrow skull, as well as elongated upper and sometimes lower canine teeth and incisors which were likely used as slashing and stabbing weapons. Postcanine teeth are generally reduced or absent. For hunting large prey, they possibly used a bite-and-retreat tactic, ambushing and taking a debilitating bite out of the target, and following it at a safe distance before its injuries exhausted it, whereupon the gorgonopsian would grapple the animal and deliver a killing bite. They would have had an exorbitant gape, possibly in excess of 90°, without having to unhinge the jaw. They markedly increased in size as time went on, growing from small skull lengths of 10–15 cm (4–6 in) in the Middle Permian to bear-like proportions of up to 60 cm (2 ft) in the Upper Permian. The latest gorgonopsians, Rubidgeinae, were the most robust of the group and could produce especially powerful bites. Gorgonopsians are thought to have been completely terrestrial and could walk with a semi-erect gait, with a similar terrestrial locomotory range as modern crocodilians. They may have been more agile than their prey items, but were probably inertial homeotherms rather than endotherms unlike contemporary therocephalians and cynodonts, and thus were probably comparatively less active. Though gorgonopsians were able to maintain a rather high body temperature, it is unclear if they would have also had sweat glands or fur (and by extension whiskers and related structures). Their brains were reminiscent of modern reptilian brains, rather than those of living mammals. Most species may have been predominantly diurnal (active during the day) though some could have been crepuscular (active at dawn or dusk) or nocturnal (active at night). They are thought to have had binocular vision, a parietal eye (which detects sunlight and maintains circadian rhythm), a keen sense of smell, a functional vomeronasal organ ("Jacobson's organ"), and possibly a rudimentary eardrum. The major therapsid groups had all evolved by 275 million years ago from a "pelycosaur" ancestor (a poorly defined group including all synapsids which are not therapsids). The therapsid takeover from pelycosaurs took place by the Middle Permian as the world progressively became drier. Gorgonopsians rose to become apex predators of their environments following the Capitanian mass extinction event which killed off the dinocephalians and some large therocephalians after the Middle Permian. Despite the existence of a single continent during the Permian, Pangaea, gorgonopsians have only been found in the Karoo Supergroup (primarily in South Africa, but also in Tanzania, Zambia, and Malawi), the Moradi Formation of Niger, and western Russia, with probable remains known from the Kundaram Formation of India. These places were semi-arid areas with highly seasonal rainfall. Gorgonopsian genera vary very little, and consequently, many species have been named based on flimsy and likely age-related differences since their discovery in the late 19th century, and the group has been subject to several taxonomic revisions. They became extinct during a phase of the Permian–Triassic extinction event taking place at the very end of the Permian, in which major volcanic activity (which would produce the Siberian Traps) and resultant massive spike in greenhouse gases caused rapid aridification due to temperature spike, acid rain, frequent wildfires, and potential breakdown of the ozone layer. The large predatory niches would be taken over by the archosaurs (namely crocodilians and dinosaurs) in the Mesozoic.
  • 1.2K
  • 17 Oct 2022
Topic Review
Struthiosaurinae
Struthiosaurinae is a subfamily of ankylosaurian dinosaurs from the Cretaceous of Europe. It is defined as "the most inclusive clade containing Europelta but not the North American Cedarpelta, Peloroplites, Sauropelta or Edmontonia" while being reinstated for a newly recognized clade of basal nodosaurids. Struthiosaurinae appeared at about exactly the same time as the North American subfamily Nodosaurinae. Struthiosaurines range all across the Cretaceous, the oldest genus being Europelta at an age of 112 Ma and the youngest being Struthiosaurus at about 85–66 Ma. It was originally mentioned by Franz Nopcsa in 1923 as a subfamily of Acanthopholidae, along with the previously defined Acanthopholinae. The family has gone through many taxonomic revisions since it was defined by Nopcsa in 1902. It is now recognized as a junior synonym of the family Nodosauridae. The subfamily now includes the genera Anoplosaurus, Europelta, Hungarosaurus, and Struthiosaurus, designated as the type genus. Because of the instability of Acanthopholis, the generic namesake of Acanthopholinae, and its current identification as a nomen dubium, Struthiosaurinae, the next named group, was decidedly used over the older one. A review of ankylosaur osteoderms was published in 2000, and reviewed the armour of Struthiosaurinae. The group was represented by the single genus Struthiosaurus, known from head, cervical, dorsal, sacral, and caudal scutes. Only a few head osteoderms were identified, so it is unknown how much of the skull was armoured. Many cervical and dorsal scutes have been preserved alongside species of Struthiosaurus. They include cervical bands, which are groups of osteoderms fused together and attached to the vertebrae, and large spines found on the shoulders of nodosaurids like Sauropelta and Edmontonia, although it is not known if the spines were fused like the later of separate like the former. It is quite possible that small ovoid scutes found on Struthiosaurus could have formed a pelvic shield like polacanthids. The caudal scutes of struthiosaurines are small and rough. Even though osteoderms are well-known, it is not certain where they were positioned on the body.
  • 904
  • 28 Nov 2022
Topic Review
Spinophorosaurus
Spinophorosaurus is a genus of sauropod dinosaur that lived in what is now Niger during the Middle Jurassic period. The first two specimens were excavated in the 2000s by German and Spanish teams under difficult conditions. The skeletons were brought to Europe and digitally replicated, making Spinophorosaurus the first sauropod to have its skeleton 3D printed, and were to be returned to Niger in the future. Together, the two specimens represented most of the skeleton of the genus, and one of the most completely known basal sauropods of its time and place. The first skeleton was made the holotype specimen of the new genus and species Spinophorosaurus nigerensis in 2009; the generic name ("spine-bearing lizard") refers to what was initially thought to be spiked osteoderms, and the specific name (Niger and -ensis) refers to where it was found. A juvenile sauropod from the same area was later assigned to the genus. The subadult holotype specimen is estimated to have been around 13 m (43 ft) in length, whereas the paratype was about 14 m (46 ft) long. The shoulder height reached by these individuals was estimated at around 4 m (13 ft), and the weight at about 7 metric tons (7.7 short tons). The braincase was short, deep, and broad, and the neuroanatomy was in some ways intermediate between that of basal sauropodomorphs and the more derived neosauropods. The teeth were spatulate (spoon shaped) and had large spaced denticles at the top of the crown, an ancestral feature in sauropods. The neck of Spinophorosaurus is one of the most completely known among sauropods, containing 13 vertebrae. The dorsal vertebrae had multiple small air-filled internal chambers, a feature typical of later more-derived sauropods. The tail was powered by strong musculature and had a rear section that was rather rigid due to long and overlapping chevron bones. Bones originally thought to be osteoderms bearing spikes placed on the tail tip were later suggested to be clavicles. Spinophorosaurus has been classified as either a very basal sauropod, or inside Eusauropoda, a more derived group. The anatomy, age, and location of specimens indicate that important developments in sauropod evolution may have occurred in North Africa, possibly controlled by climatic zones and plant biogeography. Features of the vestibular apparatus suggest that vision and coordinated eye, head, and neck movements were important in Spinophorosaurus. 3D models of the skeleton have been used to test its range of motion. One study suggests it may have been a high browser, and another examined possible mating postures. Sutures between the neural arches with the centra of the vertebrae were more complex in the front part of the trunk of Spinophorosaurus, since stresses were probably greatest in that region. Spinophorosaurus is known from the Irhazer Shale, a geological formation thought to be Middle Jurassic in age. It was formed by deposits from rivers and lakes in a great river-valley system.
  • 902
  • 24 Nov 2022
Topic Review
Antler Orogeny
The Antler orogeny was a tectonic event that began in the early Late Devonian with widespread effects continuing into the Mississippian and early Pennsylvanian. Most of the evidence for this event is in Nevada but the limits of its reach are unknown. A great volume of conglomeratic deposits of mainly Mississippian age in Nevada and adjacent areas testifies to the existence of an important tectonic event, and implies nearby areas of uplift and erosion, but the nature and cause of that event are uncertain and in dispute. Although it is known as an orogeny (mountain building event), some of the classic features of orogeny as commonly defined such as metamorphism, and granitic intrusives have not been linked to it. In spite of this, the event is universally designated as an orogeny and that practice is continued here. This article outlines what is known and unknown about the Antler orogeny and describes three current theories regarding its nature and origin.
  • 856
  • 30 Nov 2022
Topic Review
Marasuchus
Marasuchus (meaning "Mara crocodile") is a genus of basal dinosauriform archosaur which is possibly synonymous with Lagosuchus. Both genera lived during the Late Triassic in what is now La Rioja Province, Argentina . Marasuchus contains a single species, Marasuchus lilloensis. Marasuchus lilloensis was originally designated as Lagosuchus lilloensis in 1972. It was considered a new species of Lagosuchus, a contemporary archosaur described the previous year. However, a 1994 study argued that the original material of Lagosuchus was undiagnostic. This would mean that Lagosuchus and its original species (Lagosuchus talampayaensis) could be considered nomen dubia. Specimens of Lagosuchus stored at a museum in San Miguel de Tucuman were considered to be more diagnostic than those of L. talampayensis, and thus they were given a new genus: Marasuchus. A 2019 study redescribed the original material of Lagosuchus and concluded that it was valid and not readily distinguishable from Marasuchus lilloensis. This suggests that Marasuchus lilloensis is a junior synonym of Lagosuchus talampayensis. Specimens referred to the genus Marasuchus possessed some, but not all of the adaptations which traditionally characterized dinosaurs. For example, its proportions indicate that it was likely bipedal as in early dinosaurs. Also, it shared certain specific characteristics with that group, most relating to the hip and the head of the femur. Nevertheless, it lacked certain dinosaur-like features such as a perforated acetabulum, and it had several plesiomorphic ("primitive") features of the ankle.
  • 816
  • 23 Nov 2022
Topic Review
Maastrichtian Vertebrates of Lbero-Armorica
The South-Pyrenean Basin (northeastern Spain) has yielded a rich and diverse record of Upper Cretaceous (uppermost Campanian−uppermost Maastrichtian) vertebrate fossils, including the remains of some of the last European dinosaurs prior to the Cretaceous-Paleogene (K-Pg) extinction event. In this work, we update and characterize the vertebrate fossil record of the Arén Sandstone and Tremp formations in the Western Tremp Syncline, which is located in the Aragonese area of the Southern Pyrenees. The transitional and continental successions of these sedimentary units are dated to the late Maastrichtian, and exploration of their outcrops has led to the discovery of numerous fossil remains (bones, eggshells, and tracks) of dinosaurs, including hadrosauroids, sauropods, and theropods, along with other tetrapods such as crocodylomorphs, testudines, pterosaurs, squamates, and amphibians. In particular, this fossil record contains some of the youngest lambeosaurine hadrosaurids (Arenysaurus and Blasisaurus) and Mesozoic crocodylomorphs (Arenysuchus and Agaresuchus subjuniperus) in Europe, complementing the lower Maastrichtian fossil sites of the Eastern Tremp Syncline.
  • 792
  • 10 May 2021
Topic Review
“Elephant-Equus” Event
The dispersal of primitive elephantines and monodactyl equids in Eurasia has long been regarded as representative of a substantial turnover in mammal faunas, denoting the spread of open environments linked to the onset of cold and dry conditions in the Northern Hemisphere. During the 1980s, this event was named the “Elephant-Equus event” and it was correlated with the Gauss-Matuyama reversal, today corresponding to the Pliocene-Pleistocene transition and the beginning of the Quaternary, dated at ~2.6 Ma. Therefore, the Elephant-Equus event became a concept of prominent biochronological and paleoecological significance, especially in western Europe. Yet, uncertainties surrounding the taxonomy and chronology of early “elephant” and “Equus”, as well as conceptual differences in adopting (or understanding) the Elephant-Equus event as an intercontinental dispersal event or as a stratigraphic datum, engendered ambiguity and debate.
  • 777
  • 01 Jun 2023
Topic Review
Pelecanimimus
Pelecanimimus (meaning "pelican mimic") is an extinct genus of basal ("primitive") ornithomimosaurian dinosaur from the Early Cretaceous of Spain . It is notable for possessing more teeth than any other member of the Ornithomimosauria (or any other theropod), most of which were toothless.
  • 727
  • 22 Nov 2022
Topic Review
Sulphur Mountain Formation
The Sulphur Mountain Formation is a geologic formation of Early to Middle Triassic age. It is present on the western edge of the Western Canada Sedimentary Basin in the foothills and Rocky Mountains of western Alberta and northeastern British Columbia. It includes marine fossils from the time shortly after the Permian-Triassic extinction event. The Sulphur Mountain Formation was first described as a member of the Spray River Formation by P.S. Warren in 1945, who named it for Sulphur Mountain in Banff National Park. It was later raised to formation status. Its type section is located in the Spray River gorge at the southern end of Sulphur Mountain.
  • 665
  • 30 Nov 2022
Topic Review
Hosselkus Limestone
The Hosselkus Limestone is an Upper Triassic fossiliferous marine micritic limestone formation that outcrops in Plumas and Shasta Counties, California . It is known for its invertebrate fauna, most notably the many species of shelled cephalopods.
  • 572
  • 29 Sep 2022
Topic Review
Skeletal Transformations and the Origin of Baleen Whales
Mysticeti or baleen whales are a group of cetaceans including a few extant species in only six genera. Despite their scarce diversity, baleen whales represent key species in the trophic webs of the oceans as they are able to sequester enormous quantities of carbon and to disseminate nutrients that support large planktonic blooms. The study of their evolution is thus crucial to understand how and when the ecological characteristics of today’s oceans came to be.
  • 528
  • 06 Apr 2022
Topic Review
Coon Creek Formation
The Coon Creek Formation is a geologic formation located in western Tennessee and extreme northeast Mississippi. It is a sedimentary sandy marl deposit, Late Cretaceous (Maastrichtian) in age, about 70 million years old. The formation is known for producing mosasaurs and plesiosaurs, particularly at Coon Creek in McNairy County, Tennessee, which the formation is named for. Additionally, the formation produces many other marine invertebrates such as Turritella and the state fossil of Tennessee, the bivalve Pterotrigonia thoracica, as well as other fossils such as crabs.
  • 521
  • 13 Nov 2022
Topic Review
Burgess Shale Type Preservation
The Burgess Shale of British Columbia is famous for its exceptional preservation of mid-Cambrian organisms. Around 40 other sites have been discovered of a similar age, with soft tissues preserved in a similar, though not identical, fashion. Additional sites with a similar form of preservation are known from the Ediacaran and Ordovician periods. These various shales are of great importance in the reconstruction of the ecosystems immediately after the Cambrian explosion. The taphonomic regime results in soft tissue being preserved, which means that organisms without hard parts that could be conventionally fossilised can be seen; also, we gain an insight into the organs of more familiar organisms such as the trilobites. The most famous localities preserving organisms in this fashion are the Canadian Burgess Shale, the Chinese Chengjiang fauna, and the more remote Sirius Passet in north Greenland. However, a number of other localities also exist.
  • 518
  • 28 Oct 2022
Topic Review
Kirtlandian
The Kirtlandian is a North American land-vertebrate faunal age of the Cretaceous period, following the Judithian and succeeded by the Edmontonian. It lasted about 2 million years, ca 74.8 to 72.8 Mya and is characterized by the ceratopsian Pentaceratops sternbergii, which lived throughout the Kirtlandian. It was first named by R.M. Sullivan and S.G. Lucas in 2003 as a faunal age for the Kirtland and Fruitland formations. Previously, only five land-vertebrate ages were identified from the Late Cretaceous. as identified by Loris S. Russell in 1975, they include the Paluxian, Aquilan, Judithian, Edmontonian, and the Lancian. Before the naming of the Kirtlandian, three gaps, between the Paluxian and Aquilan, the Aquilan and the Judithian, and the Judithian and Edmontonian, were identified but not named. The Fruitland Formation measures 97 to 107 metres (318 to 351 ft) thick, and with the 594 metres (1,949 ft) of the Kirtland Formation, the Kirtlandian consists of 701 metres (2,300 ft) of sediments. The rock types within the formations are primarily coal beds, but also include sandstone, siltstone, mudstone, and shale. Within the sediments with a Kirtlandian age, two local faunas, the Hunter Wash local fauna, and the Willow Wash local fauna, have been identified. The currently accepted date of the Kirtlandian is 74.8 to 72.8 million years ago.
  • 441
  • 04 Nov 2022
Topic Review
2017 in Paleomalacology
This list, 2017 in paleomalacology, is a list of new taxa of ammonites and other fossil cephalopods, as well as fossil gastropods, bivalves and other molluscs that are scheduled to be described during the year 2017, as well as other significant discoveries and events related to molluscan paleontology that are scheduled to occur in the year 2017.
  • 421
  • 27 Nov 2022
Topic Review
Large Mammals as Vitamin C Sources for MIS 3 Hominins
The acquisition of large prey by hominins living during the Marine Isotope Stage 3, including Neanderthals and Anatomically Modern Humans, had nutritional and bioenergetic implications: these contain high fat amounts, provide a high energy return, and the strategies and skills required to acquire small prey were different from those required to acquire the former. Vitamin C availability at several MIS 3 periods could have had a strong seasonal variability and would have been decisive for hominin groups’ survival. 
  • 313
  • 19 Jan 2024
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