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HandWiki. List of Cyphophthalmi Species. Encyclopedia. Available online: https://encyclopedia.pub/entry/28514 (accessed on 20 December 2025).
HandWiki. List of Cyphophthalmi Species. Encyclopedia. Available at: https://encyclopedia.pub/entry/28514. Accessed December 20, 2025.
HandWiki. "List of Cyphophthalmi Species" Encyclopedia, https://encyclopedia.pub/entry/28514 (accessed December 20, 2025).
HandWiki. (2022, October 09). List of Cyphophthalmi Species. In Encyclopedia. https://encyclopedia.pub/entry/28514
HandWiki. "List of Cyphophthalmi Species." Encyclopedia. Web. 09 October, 2022.
List of Cyphophthalmi Species
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The content is sourced from: https://handwiki.org/wiki/Biology:List_of_Cyphophthalmi_species

This page lists all described species of the harvestman suborder Cyphophthalmi. Unless otherwise noted, information is taken from Giribet's 2000 taxonomic catalogue and from Giribet et al., 2012.

cyphophthalmi harvestman suborder

1. Boreophthalmi

1.1. Sironidae

  • Cyphophthalmus[1]
  • Cyphophthalmus beschkovi (Mitov, 1994)
  • Cyphophthalmus bithynicus (Gruber, 1969)
  • Cyphophthalmus conocephalus Karaman, 2009
  • Cyphophthalmus corfuanus (Kratochvíl, 1938)
  • Cyphophthalmus duricorius Joseph, 1868
  • Cyphophthalmus eratoae (Juberthie, 1968)
  • Cyphophthalmus ere Karaman, 2008
  • Cyphophthalmus gjorgjevici (Hadži, 1933)
  • Cyphophthalmus gordani Karaman, 2009
  • Cyphophthalmus hlavaci Karaman, 2009
  • Cyphophthalmus klisurae (Hadži, 1973)
  • Cyphophthalmus kratochvili Karaman, 2009
  • Cyphophthalmus markoi Karaman, 2008
  • Cyphophthalmus martensi Karaman, 2009
  • Cyphophthalmus minutus (Kratochvíl, 1938)
  • Cyphophthalmus neretvanus Karaman, 2009
  • Cyphophthalmus noctiphilus (Kratochvíl, 1940)
  • Cyphophthalmus nonveilleri Karaman, 2008
  • Cyphophthalmus ognjenovici Karaman, 2009
  • Cyphophthalmus ohridanus (Hadži, 1973)
  • Cyphophthalmus paradoxus (Kratochvíl, 1958)
  • Cyphophthalmus paragamiani Karaman, 2009
  • Cyphophthalmus rumijae Karaman, 2009
  • Cyphophthalmus serbicus (Hadži, 1973)
  • Cyphophthalmus silhavyi (Kratochvíl, 1938)
  • Cyphophthalmus solentiensis Dreszer, Raða & Giribet, 2015[2]
  • Cyphophthalmus teyrovskyi (Kratochvíl, 1938)
  • Cyphophthalmus thracicus Karaman, 2009
  • Cyphophthalmus trebinjanus Karaman, 2009
  • Cyphophthalmus yalovensis (Gruber, 1969)
  • Cyphophthalmus zetae Karaman, 2009
  • Iberosiro de Bivort & Giribet, 2004[3]
  • Iberosiro distylos de Bivort & Giribet, 2009
  • Iberosiro rosae Giribet, Merino-Sáinz & Benavides, 2017
  • Marwe Shear, 1985
  • Marwe coarctata Shear, 1985
  • Odontosiro Juberthie, 1961
  • Odontosiro lusitanicus Juberthie, 1961
  • Paramiopsalis Juberthie, 1962[3]
  • Paramiopsalis anadonae Giribet, Merino-Sáinz & Benavides, 2017
  • Paramiopsalis eduardoi Murienne & Giribet, 2009
  • Paramiopsalis ramblae Benavides & Giribet, 2017
  • Paramiopsalis ramulosus Juberthie, 1962
  • Parasiro Hansen and Sørensen, 1904
  • Parasiro coiffaiti Juberthie, 1956
  • Parasiro corsicus (Simon, 1872)
  • Parasiro minor Juberthie, 1958
  • Siro Latreille, 1796[3]
  • Siro acaroides (Ewing, 1923)
  • Siro boyerae Giribet & Shear, 2010
  • Siro calaveras Giribet & Shear, 2010
  • Siro carpaticus Rafalski, 1956
  • Siro clousi Giribet & Shear, 2010
  • Siro crassus Novak & Giribet, 2006
  • Siro exilis Hoffman, 1963
  • Siro kamiakensis (Newell, 1943)
  • Siro ligiae Giribet, 2017
  • Siro rubens Latreille, 1804
  • Siro richarti Benavides & Giribet, 2017
  • Siro shasta Giribet & Shear, 2010
  • Siro sonoma Shear, 1980
  • Siro valleorum Chemini, 1990
  • Suzukielus Juberthie, 1970
  • Suzukielus sauteri (Roewer, 1916)

1.2. Stylocellidae[4]

Fangensinae

  • Fangensis Rambla, 1994
  • Fangensis cavernarus Schwendinger & Giribet, 2005
  • Fangensis leclerci Rambla, 1994
  • Fangensis spelaeus Schwendinger & Giribet, 2005
  • Giribetia Clouse, 2012
  • Giribetia insulanus (Schwendinger & Giribet, 2005)

Leptopsalidinae[5]

  • Leptopsalis Thorell, 1882
  • Leptopsalis beccarii Thorell, 1882
  • Leptopsalis dumoga (Shear, 1993)
  • Leptopsalis hillyardi (Shear, 1993)
  • Leptopsalis javana (Thorell, 1882)
  • Leptopsalis laevichelis (Roewer, 1942)
  • Leptopsalis lydekkeri (Clouse & Giribet, 2007)
  • Leptopsalis modesta (Hansen & Sørensen, 1904)
  • Leptopsalis novaguinea (Clouse & Giribet, 2007)
  • Leptopsalis pangrango (Shear, 1993)
  • Leptopsalis ramblae (Giribet, 2002)
  • Leptopsalis sedgwicki (Shear, 1979)
  • Leptopsalis sulcata (Hansen & Sørensen, 1904)
  • Leptopsalis tambusisi (Shear, 1993)
  • Leptopsalis thorellii (Hansen & Sørensen, 1904)
  • Leptopsalis weberii (Hansen & Sørensen, 1904)
  • Miopsalis Thorell, 1890
  • Miopsalis collinsi (Shear, 1993)
  • Miopsalis globosa (Schwendinger & Giribet, 2004)
  • Miopsalis gryllospeca (Shear, 1993)
  • Miopsalis kinabalu (Shear, 1993)
  • Miopsalis leakeyi (Shear, 1993)
  • Miopsalis lionota (Pocock, 1897)
  • Miopsalis mulu (Shear, 1993)
  • Miopsalis pocockii (Hansen & Sørensen, 1904)
  • Miopsalis pulicaria Thorell, 1890
  • Miopsalis sabah (Shear, 1993)
  • Miopsalis silhavyi (Rambla, 1991)
  • Miopsalis tarumpitao (Shear, 1993)

Stylocellinae

  • Meghalaya Giribet, Sharma, & Bastawade, 2007
  • Meghalaya annandalei Giribet, Sharma, & Bastawade, 2007
  • Stylocellus Westwood, 1874
  • Stylocellus lornei Clouse, 2012
  • Stylocellus sumatranus Westwood, 1874

2. Scopulophthalmi

2.1. Pettalidae

  • Aoraki Boyer & Giribet, 2007[6]
  • Aoraki calcarobtusa (Forster, 1952)
  • Aoraki calcarobtusa calcarobtusa (Forster, 1952)
  • Aoraki calcarobtusa westlandica (Forster, 1952)
  • Aoraki crypta (Forster, 1948)
  • Aoraki denticulata (Forster, 1948)
  • Aoraki denticulata denticulata (Forster, 1948)
  • Aoraki denticulata major (Forster, 1948)
  • Aoraki granulosa (Forster, 1952)
  • Aoraki healyi (Forster, 1948)
  • Aoraki inerma (Forster, 1948)
  • Aoraki inerma inerma (Forster, 1948)
  • Aoraki inerma stephenesis (Forster, 1952)
  • Aoraki longitarsa (Forster, 1952)
  • Aoraki tumidata (Forster, 1948)
  • Austropurcellia Juberthie, 1988[7]
  • Austropurcellia absens Boyer & Popkin-Hall, 2015
  • Austropurcellia acuta Popkin-Hall & Boyer, 2014
  • Austropurcellia alata Boyer & Reuter, 2012
  • Austropurcellia arcticosa (Cantrell, 1980)
  • Austropurcellia barbata Popkin-Hall & Boyer, 2014
  • Austropurcellia cadens Baker & Boyer, 2015
  • Austropurcellia capricornia (Davies, 1977)
  • Austropurcellia clousei Boyer, Baker & Popkin-Hall, 2015
  • Austropurcellia culminis Boyer & Reuter, 2012
  • Austropurcellia daviesae (Juberthie, 1989)
  • Austropurcellis despectata Boyer & Reuter, 2012
  • Austropurcellia finniganensis Popkin-Hall, Jay & Boyer, 2016
  • Austropurcellia forsteri (Juberthie, 2000)
  • Austropurcellia fragosa Popkin-Hall, Jay & Boyer, 2016
  • Austropurcellia giribeti Boyer & Quay, 2015
  • Austropurcellia megatanka Jay, Coblens & Boyer, 2016
  • Austropurcellia monteithi Jay, Popkin-Hall, Coblens & Boyer, 2016
  • Austropurcellia riedeli Jay, Oberski & Boyer, 2016
  • Austropurcellia nuda Popkin-Hall, Jay & Boyer, 2016
  • Austropurcellia scoparia Juberthie, 1988
  • Austropurcellia sharmai Boyer & Quay, 2015
  • Austropurcellia superbensis Popkin-Hall & Boyer, 2014
  • Austropurcellia tholei Baker & Boyer, 2015
  • Austropurcellia vicina Boyer & Reuter, 2012
  • Austropurcellia woodwardi (Forster, 1955)
  • Chileogovea Roewer, 1961
  • Chileogovea jocasta Shear, 1993
  • Chileogovea oedipus Roewer, 1961
  • Karripurcellia Giribet, 2003[8]
  • Karripurcellia harveyi Giribet, 2003
  • Karripurcellia peckorum Giribet, 2003
  • Karripurcellia sierwaldae Giribet, 2003
  • Manangotria Shear & Gruber, 1996
  • Manangotria taolanaro Shear & Gruber, 1996
  • Neopurcellia Forster, 1948[6]
  • Neopurcellia salmoni Forster, 1948
  • Parapurcellia Rosas Costa, 1950[9]
  • Parapurcellia amatola de Bivort & Giribet, 2010
  • Parapurcellia convexa de Bivort & Giribet, 2010
  • Parapurcellia fissa (Lawrence, 1939)
  • Parapurcellia minuta de Bivort & Giribet, 2010
  • Parapurcellia monticola (Lawrence, 1939)
  • Parapurcellia natalia de Bivort & Giribet, 2010
  • Parapurcellia rumpiana (Lawrence, 1933)
  • Parapurcellia peregrinator (Lawrence, 1963)
  • Parapurcellia silvicola (Lawrence, 1939)
  • Parapurcellia staregai de Bivort & Giribet, 2010
  • Parapurcellia transvaalica (Lawrence, 1963)[10]
  • Pettalus Thorell, 1876
  • Pettalus brevicauda Pocock, 1897
  • Pettalus cimiciformis (Cambridge, 1875)
  • Pettalus lampetides Sharma & Giribet 2006[11]
  • Pettalus thwaitesi Sharma, Karunarathna & Giribet, 2009[12]
  • Purcellia Hansen & Sørensen, 1904[9]
  • Purcellia argasiformis (Lawrence, 1931)[10]
  • Purcellia griswaldi de Bivort & Giribet, 2010
  • Purcellia illustrans Hansen and Sørensen, 1904
  • Purcellia lawrencei de Bivort & Giribet, 2010
  • Purcellia leleupi Staręga, 2008[13]
  • Rakaia Hirst, 1925[6]
  • Rakaia antipodiana Hirst, 1925
  • Rakaia collaria Roewer, 1942
  • Rakaia dorothea Phillips and Grimmett, 1932
  • Rakaia florensis (Forster, 1948)
  • Rakaia isolata Forster, 1952
  • Rakaia lindsayi Forster, 1952
  • Rakaia macra Boyer and Giribet, 2003
  • Rakaia magna Forster, 1948
  • Rakaia magna australis Forster, 1952
  • Rakaia magna magna Forster, 1948
  • Rakaia media Forster, 1948
  • Rakaia media media Forster, 1948
  • Rakaia media insula Forster, 1952
  • Rakaia minutissima (Forster, 1948)
  • Rakaia pauli Forster, 1952
  • Rakaia solitaria Forster, 1948
  • Rakaia sorenseni Forster, 1952
  • Rakaia sorenseni digitata Forster, 1952
  • Rakaia sorenseni sorenseni Forster, 1952
  • Rakaia stewartiensis Forster, 1948
  • Rakaia uniloca Forster, 1952

3. Sternophthalmi

3.1. Neogoveidae[14]

  • Brasiliogovea Martens, 1969
  • Brasiliogovea aphantostylus Benavides, Hormiga, & Giribet, 2019
  • Brasiliogovea chiribiqueta Benavides & Giribet, 2013
  • Brasiliogovea microphaga Martens, 1969
  • Brasiliogovea microstylus Benavides, Hormiga, & Giribet, 2019
  • Brasiliogovea yacambuensis Benavides, Hormiga, & Giribet, 2019
  • Canga DaSilva, Pinto-da-Rocha & Giribet, 2010
  • Canga renatae DaSilva, Pinto-da-Rocha & Giribet, 2010
  • Huitaca Shear, 1979
  • Huitaca boyacanensis Benavides & Giribet, 2013
  • Huitaca bitaco Benavides & Giribet, 2013
  • Huitaca caldas Benavides & Giribet, 2013
  • Huitaca depressa Benavides & Giribet, 2013
  • Huitaca sharkeyi Benavides & Giribet, 2013
  • Huitaca tama Benavides & Giribet, 2013
  • Huitaca ventralis Shear, 1979
  • Leggogovia Benavides & Giribet, 2019
  • Leggogovia pabsgarnoni (Legg. 1990)
  • Metagovea Rosas-Costa, 1950
  • Metagovea disparunguis Rosas Costa, 1950
  • Metagovea ligiae Giupponi & Kury, 2015
  • Metagovea matapi Benavides, Hormiga, & Giribet, 2019
  • Metagovea philipi Goodnight & Goodnight, 1980
  • Metagovea planada Benavides, Hormiga, & Giribet, 2019
  • Metasiro Juberthie, 1960[15]
  • Metasiro americanus (Davis, 1933)
  • Metasiro savannahensis Clouse & Wheeler, 2014
  • Metasiro sassafrasensis Clouse & Wheeler, 2014
  • Microgovia Benavides, Hormiga, & Giribet, 2019
  • Microgovia chenepau Benavides, Hormiga, & Giribet, 2019
  • Microgovia oviformis Martens, 1969
  • Neogovea Hinton, 1938
  • Neogovea branstetteri Benavides, Hormiga, & Giribet, 2019
  • Neogovea enigmatica Martens, 2019
  • Neogovea hormigai Benavides & Giribet, 2013
  • Neogovea immsi Hinton, 1938
  • Neogovea kamakusa Shear, 1977
  • Neogovea kartabo Shear, 1977
  • Neogovea matawai Benavides, Hormiga, & Giribet, 2019
  • Neogovea virginie Jocqué & Jocqué, 2011
  • Parogovia Hansen, 1921
  • Parogovia gabonica (Juberthie, 1969)
  • Parogovia montealensis Benavides & Giribet, 2019
  • Parogovia parasironoides Hiřman, Kotyk, Kotyková Varadínová,& Šťáhlavský, 2018
  • Parogovia prietoi Benavides & Giribet, 2019
  • Parogovia putnami Benavides & Giribet, 2019
  • Parogovia sironoides Hansen, 1921
  • Tucanogovea Karaman, 2014
  • Tucanogovea schusteri Karaman, 2014
  • Waiwaigovia Benavides, Hormiga, & Giribet, 2019
  • Waiwaigovia shultzi Benavides, Hormiga, & Giribet, 2019

3.2. Ogoveidae[16]

  • Ogovea Roewer, 1923
  • Ogovea cameroonensis Giribet & Prieto, 2003
  • Ogovea grossa (Hansen & Sørensen, 1904)
  • Ogovea nasuta (Hansen, 1921)

3.3. Troglosironidae[17]

  • Troglosiro Juberthie, 1979
  • Troglosiro aelleni Juberthie, 1979
  • Troglosiro brevifossa Sharma & Giribet, 2009
  • Troglosiro juberthiei Shear, 1993
  • Troglosiro longifossa Sharma & Giribet, 2005
  • Troglosiro monteithi Sharma & Giribet, 2009
  • Troglosiro ninqua Shear, 1993
  • Troglosiro oscitatio Sharma & Giribet, 2009
  • Troglosiro platnicki Shear, 1993
  • Troglosiro raveni Shear, 1993
  • Troglosiro sheari Sharma & Giribet, 2009
  • Troglosiro tillierorum Shear, 1993
  • Troglosiro urbanus Sharma & Giribet, 2009
  • Troglosiro wilsoni Sharma & Giribet, 2009

4. incertae Cedis

  • Ankaratra Shear & Gruber, 1996
  • Ankaratra franzi Shear & Gruber, 1996
  • Shearogovea Giribet, 2011
  • Shearogovea mexasca (Shear, 1977)

References

  1. Karaman, Ivo M. (2009-06-01). "The taxonomical status and diversity of Balkan sironids (Opiliones, Cyphophthalmi) with descriptions of twelve new species" (in en). Zoological Journal of the Linnean Society 156 (2): 260–318. doi:10.1111/j.1096-3642.2009.00446.x. ISSN 0024-4082.  https://dx.doi.org/10.1111%2Fj.1096-3642.2009.00446.x
  2. Giribet, Gonzalo; Raða, Tonći; Dreszer, Taras B. (2015). "Cyphophthalmus solentiensis sp. nov. (Cyphophthalmi, Sironidae), a New Endogean Mite Harvestman Species from Croatia, with an Application of Confocal Laser Microscopy to Illustrate Genitalia in Opiliones". Breviora 543 (1): 1–15. doi:10.3099/MCZ18.1. ISSN 0006-9698.  https://dx.doi.org/10.3099%2FMCZ18.1
  3. Merino-Sáinz, Izaskun; Benavides, Ligia R.; Giribet, Gonzalo (2017-08-30). "The systematics and biogeography of the mite harvestman family Sironidae (Arachnida : Opiliones : Cyphophthalmi) with the description of five new species" (in en). Invertebrate Systematics 31 (4): 456–491. doi:10.1071/IS16086. ISSN 1447-2600.  https://dx.doi.org/10.1071%2FIS16086
  4. Clouse, Ronald (2012). "The lineages of Stylocellidae (Arachnida: Opiliones: Cyphophthalmi)". Zootaxa 3595 (3595): 1–34. doi:10.11646/zootaxa.3595.1.1. ISSN 1175-5334. http://www.museunacional.ufrj.br/mndi/Aracnologia/pdfliteratura/Cypho/Clouse%202012%20-%20The%20lineages%20of%20Stylocellidae.pdf. 
  5. Kury, Adriano (2018). "The correct spelling of Leptopsalidinae (Arachnida: Opiliones: Cyphophthalmi).". Arachnida – Rivista Aracnologica Italiana 18: 40–43. https://www.researchgate.net/publication/328830221. 
  6. Boyer, Sarah; Giribet, Gonzalo (2007). "A new model Gondwanan taxon: systematics and biogeography of the harvestman family Pettalidae (Arachnida, Opiliones, Cyphophthalmi), with a taxonomic revision of genera from Australia and New Zealand". Cladistics 23 (4): 337–361. doi:10.1111/j.1096-0031.2007.00149.x. https://boyerlab.weebly.com/uploads/8/5/9/2/85928266/boyer___giribet_2007.pdf. 
  7. Boyer, Sarah; Sharma, Prashant; Oberski, Jill; Coblens, Michelle; Popkin-Hall, Zachary; Jay, Katya (2016-04-05). "New species of Austropurcellia, cryptic short-range endemic mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) from Australia's Wet Tropics biodiversity hotspot" (in en). ZooKeys (586): 37–93. doi:10.3897/zookeys.586.6774. ISSN 1313-2970. PMID 27199608. https://zookeys.pensoft.net/article/6774/. 
  8. Giribet, Gonzalo (2003). "Karripurcellia, a new pettalid genus (Arachnida: Opiliones: Cyphophthalmi) from Western Australia, with a cladistic analysis of the family Pettalidae". Invertebrate Systematics 17 (3): 387–406. doi:10.1071/IS02014.  https://dx.doi.org/10.1071%2FIS02014
  9. de Bivort, Benjamin; Giribet, Gonzalo (2010). "A systematic revision of the South African Pettalidae (Arachnida:Opiliones:Cyphophthalmi) based on a combined analysis of discrete and continuous morphological characters with the description of seven new species". Invertebrate Systematics 24 (4): 371–406. doi:10.1071/IS10015. https://pdfs.semanticscholar.org/7d11/88f81d4e9920293581a68e6f1c14f85555c3.pdf?_ga=2.149457398.1725305608.1565278076-2000852403.1565278076. 
  10. Giribet, Gonzalo; Boyer, Sarah; Baker, Caitlin; Fernandéz, Rosa; Sharma, Prashant; de Bivort, Benjamin; Daniels, Savel; Harvey, Mark et al. (2016). "A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling". Zoological Journal of the Linnean Society 178 (3): 523–545. doi:10.1111/zoj.12419. https://www.macalester.edu/~boyer/site/Publications_files/Giribet%20et%20al%202016.pdf. 
  11. Sharma, Prashant; Giribet, Gonzalo (2006). "A new Pettalus species (Opiliones, Cyphophthalmi, Pettalidae) from Sri Lanka with a discussion on the evolution of eyes in Cyphophthalmi". The Journal of Arachnology 34 (2): 331–341. doi:10.1636/H05-33.1. http://www.americanarachnology.org/JoA_free/JoA_v34_n2/arac-34-2-331.pdf. 
  12. Sharma, Prashant; Karunarathna, Indika; Giribet, Gonzalo (2009). "On the endemic Sri Lankan genus Pettalus (Opiliones, Cyphophthalmi, Pettalidae) with a description of a new species and a discussion of its diversity". Journal of Arachnology 37 (1): 60–7. doi:10.1636/H08-56.1. http://www.americanarachnology.org/JoA_free/JoA_v37_n1/arac-37-01-60.pdf. 
  13. Staręga, Wojciech (2008). "The second species of Purcellia HANSEN et SØRENSEN, 1904 (Arachnida: Opiliones: Pettalidae) from South Africa". Polish Journal of Entomology 77 (1): 51–56. http://pte.au.poznan.pl/ppe/PJE_2008/06_Starega.pdf. 
  14. Benavides, Ligia; Hormiga, Gustavo; Giribet, Gonzalo (2019). "Phylogeny, evolution and systematic revision of the mite harvestman family Neogoveidae (Opiliones Cyphophthalmi)". Invertebrate Systematics 33 (1): 101–180. doi:10.1071/IS18018. https://www.researchgate.net/publication/327040072. 
  15. Clouse, Ronald; Wheeler, Ward (2014). "Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates". Zootaxa 3814 (2): 177–201. doi:10.11646/zootaxa.3814.2.2. ISSN 1175-5334. PMID 24943422. https://www.researchgate.net/publication/263291268. 
  16. Giribet, Gonzalo; Prieto, Carlos E. (2003-12-31). "A new Afrotropical Ogovea (Opiliones, Cyphophthalmi) from Cameroon, with a discussion on the taxonomic characters in the family Ogoveidae". Zootaxa 329: 1–18. doi:10.5281/zenodo.156214. https://www.researchgate.net/publication/281626631. 
  17. Sharma, Prashant; Giribet, Gonzalo (2009). "The family Troglosironidae (Opiliones: Cyphophthalmi) of New Caledonia". Zoologia Neocaledonica. 7. Mémoires du Muséum national d’Histoire naturelle. pp. 205–245. ISBN 9782856536186. https://www.researchgate.net/publication/232643583. 
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